1Department of Biology, Faculty of Sciences of Bizerte, University of Carthage, Tunisia
2UR13AGR09-Integrated Horticultural Production in the Tunisian Centre-East, Regional Research Centre on Horticulture and Organic Agriculture, University of Sousse, 4042, Chott-Mariem, Tunisia
3Laboratory of Genetics and Plant Breeding, National Agronomic Institute of Tunis, 43 Avenue Charles Nicolle, 1082 Tunis, Tunisia
4UR13AGR05-Agrobiodiversity, The Higher Agronomic Institute of Chott-Mariem, University of Sousse, 4042, Chott-Mariem, Tunisia
*Corresponding Author:
Nefzi Ahlem, Department of Biology, Faculty of Sciences of Bizerte, University of Carthage, Tunisia, Tel: +0021696532560 Email: